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Eucalyptus ancophila

Eucalyptus ancophila L.A.S.Johnson & K.D. Hill, Telopea 4: 82 (1990).

T: N.S.W., North Coast, 21 miles (33 km) NW of Kempsey, at western ridge of Mungay Creek, Feb. 1975, T.J.Evans NSW 222245; holo: NSW.

Tree to 35 m tall. Forming a lignotuber.
Grey ironbark throughout or on trunk and larger branches only, smooth bark pale grey.
Juvenile growth (coppice or field seedlings to 50 cm): stems square in cross-section; juvenile leaves petiolate, opposite for 4 to 7 nodes then alternate, ovate becoming lanceolate, discolorous, glossy, green.
Adult leaves alternate, petiolate; blade lanceolate, 9-20 cm long, 1.8-4.8 cm wide, base tapering evenly to petiole, margin entire, slightly discolorous (rarely concolorous), more or less glossy, green,
side-veins greater than 45° to midrib, densely to very densely reticulate, intramarginal vein parallel to and just within margin or well removed from it, oil glands few, mostly intersectional.
Inflorescences terminal panicles, peduncles 0.3-1 cm long; buds 7, pedicellate, ovoid to diamond-shaped, scar present, operculum beaked to conical, most stamens inflexed but usually a few irregularly flexed, with outer staminodes, anthers adnate, positioned obliquely at filament tip, cuboid, dehiscing by terminal pores, style long, locules 3 or 4, the placentae each with 4 vertical ovule rows; flowers white.
Fruit pedicellate, obconical or barrel-shaped, 0.5-0.7 cm wide, disc descending, valves 3 or 4, enclosed.
Seed brown, 1.4-2 mm long, flattened-ovoid or pointed at one end, dorsal surface shallowly reticulate, hilum ventral.

Cultivated seedlings (measured at ca node 10): cotyledons oblong; stems squared in cross-section; leaves always petiolate, opposite until nodes 4 to 7 then alternate, ovate to broadly so, 7.5-9.5 cm long, 2.5-5.5 cm wide, base tapering, discolorous, dull, green, becoming distinctly glossy after node 5.


Eucalyptus ancophila (Greek ancophila, valley lover).

A medium-sized to tall forest tree known only from the Kempsey-Bellingen districts of north-coastal New South Wales. One of a group of ironbarks distinguished by the combination of discolorous leaves, terminal inflorescences and flowers with staminodes. It has glossy green leaves which distinguish it from E. fusiformis , non-ribbed or non-angled fruit, which distinguishes it from E. tetrapleura and E. fusiformis , and longer leaves than E. placita.

Eucalyptus ancophila belongs in Eucalyptus subgenus Symphyomyrtus section Adnataria because the buds have two opercula, ovules are in four rows, seeds are flattened-ovoid, cotyledons are reniform, and anthers are rigid on the staminal filaments. Within section Adnataria, E. ancophila is part of a subgroup, series Rhodoxylon subseries Discolores being trees with ironbark, terminal inflorescences, buds that lose the outer operculum early in developement, have inflexed stamens, the outer stamens sterile (staminodes), a broad staminal ring that can often be seen on the fruit but ultimately is deciduous, and discolorous adult leaves. There are 4 other species in this subseries, E. placita , E. dorsiventralis , E. paniculata and the restricted Queensland endemic E. decolor.