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Eucalyptus angulosa Schauer in W.C. Walpers, Repert. Bot.
Syst. 2: 925 (1843).
E. cuspidata Turcz., Bull. Soc. Imp. Naturalistes Moscou
22(2): 21 (1849).
T: Western Australia: Swan River Colony, J.Drummond 4: 75;
lectotype BM; isolecto: CGE, FI, K, PERTH, W. Refer to K.D. Hill
et al., Telopea 9: 264 (2001) for further discussion.
E. incrassata var. angulosa (Schauer) Benth., Fl.
Austral. 3: 231 (1867) T: Australia, collector unknown; herbarium
of cited specimen unknown to us.
E. grossifolia L.A.S.Johnson & K.D.Hill,
Telopea 9: 271 (2001). T: Western Australia: 1.3 km from
Kambellup on Woogenellup rd (34° 34'S, 117°59'E), 12 Nov.
1986, K.D.Hill 2444, L.A.S.Johnson & D.F.Blaxell; holo:
NSW; iso: PERTH.
Mallee to 5 m tall. Forming
a lignotuber.
Bark smooth throughout with some grey, loose imperfectly
decorticated strips of rough bark accumulating on lower stems; smooth
bark pale grey, grey-brown, white or pinkish to coppery with ribbons
of decorticated bark in the upper branches.
Branchlets with or without pith oil glands.
Juvenile growth (coppice or field seedlings to 50 cm):
stems rounded or squared in cross-section; juvenile leaves alternate,
petiolate, ovate or elliptical, 4.5-9 cm long, 2.5-4 cm wide, margin
entire, green.
Adult leaves alternate, petiole 1-3 cm long; blade lanceolate
to broadly lanceolate to ovate-elliptic, 5-14 cm long, 1.7-5 cm
wide, base tapering evenly to petiole, margin entire or sometimes
shallowly and distantly denticulate, thick, concolorous, glossy,
green, side-veins greater than 45°
to midrib, moderately to very
densely reticulate, the veinlets erose, intramarginal vein well
removed from margin, oil glands numerous, intersectional.
Inflorescences axillary unbranched, peduncles flattened,
erect to spreading, 1-2.5 cm long; buds 3 or 7, pedicellate, oblong
to ovoid to pyriform, with coarse longitudinal ribs, scar present,
operculum conical to shortly beaked, stamens inflexed, anthers cuboid
to rounded, versatile, dorsifixed, dehiscing by longitudinal slits
(non-confluent), style long, locules 3 or 4, the placentae each
with 6 or 8 vertical ovule rows; flowers creamy white to pale yellow
(rarely pale pink).
Fruit pedicellate, cup-shaped, cylindrical, hemispherical
or urceolate, 1-2.5 cm wide, coarsely ribbed longitudinally, disc
descending, valves 3 or 4, enclosed.
Seed blackish grey, 2-4.5 mm long, flattened-pyramidal to
D-shaped or cuboid, with ventral ridges, dorsal surface more or
less smooth to slightly pitted, sometimes slightly lacunose, often
with a narrow marginal flange, hilum ventral.
Cultivated seedling (measured at ca node 10): cotyledons
reniform; stems squared or rounded in cross-section; leaves always
petiolate, opposite for 2 to 6 nodes then alternate, deltoid to
ovate, 4-8 cm long, 2-6 cm wide, base tapering, rounded or truncate,
dull, grey-green becoming green.
NOTES
Eucalyptus angulosa (Latin, angulosus, angled, referring
to the branchlets and buds).
A mallee of the southern Australian coast from West Cape Howe to
east of Esperance in Western Australia, and also southern Eyre Peninsula
and southern Fleurieu Peninsula in South Australia but not on Kangaroo
Island. The bark is smooth but the stems often have partly shed
strips of bark hanging. The adult leaves are thick and broadly lanceolate.
Eucalyptus angulosa belongs in Eucalyptus subgenus
Symphyomyrtus section Dumaria because the buds have
two opercula, stamens are strongly inflexed, ovules are in 6 or
8 rows on the placentae and cotyledons are reniform. E. incrassata
, E. angulosa, E. captiosa , E. singularis
and E. ceratocorys are closely related within this
section Dumaria, forming series Incrassatae.
E. angulosa is a mallee of coastal headlands and dunes with
very coarse leaves, buds and fruits. E. incrassata , with
slightly smaller buds, fruits and leaves, is sub-coastal
to inland in distribution as a component of mallee communities.
The narrow-leaved and small-budded sub-coastal mallee, E. captiosa
, and coarse-leaved inland mallee with buds that are contracted
about the middle, E. ceratocorys , both have prominently
beaked opercula. The fifth species in the series is the mallet E.
singularis which has pendulous buds on slender peduncles and
pedicels and has prominently beaked opercula.
The recently described species, E. grossifolia,
from the Kamballup area of Western Australia, south of the Stirling
Range, is not recognized here as being sufficiently distinct from
E. angulosa. It differs primarily in having a more open,
erect habit which may be due to the sub-coastal habitat, compared
to the bushy habit of more coastal-growing plants of E. angulosa.
This Kamballup form of E. angulosa is locally important for
land restoration work.
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