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Asplenium

Key modified ex Copeland 1949, Philip, J. Sci. 78:207 (incl. Diplora) and Holttum 1974, Gard. Bull. Sing. 27:145 (sect. Thamnopteris).

Species in Papuasia

1a.

Fronds simple and entire
2
Fronds pinnatifid, pinnate or more compound
18
2a. Fronds with a continuous intramarginal vein, often nest-forming
(sect. Thamopteris)

3

Veins free to the margin, fronds not nest-forming
8
3a. Sori on almost all veins, including both branches of a vein which forks near costa; indusia less than 0.5 mm wide
4
Sori usually on one branch of each vein; indusia more than 0.5 mm wide
7
4a. Fronds nest-forming
5
Fronds not nest forming
A. spirale
5a. Fronds to almost 20 cm wide; apex gradually attenuate; costa prominent and squarish above, +/- flattened below
A. nidus var. nidus
Fronds 25-25 cm wide; apex broadly rounded; costa rounded above, rounded keeled below
6
6a. Fronds to c. 50 cm long, +/- ascending
A. nidiforme
Fronds 1m or more long +/- arching
A. musifolium
7a. Fronds to 20 cm or more wide with dilated bases; indusia more than 1 mm wide
A. cymbifolium
Fronds not over 12 cm wide, bases not dilated; indusia less than 1 mm wide
A. phylitidis ssp. Malesicum
8a. Sori paired, facing one another
(Diplora)

9

Sori not paired, all facing apically
(Asplenium)

11

9a. Fronds uniform, oblanceolate
A. schizocarpa
Juvenile fronds compound, mature fronds linear-lanceolate
10
10a. Sori on most veins of fully fertile fronds
A. scolopendropsis (Diplora durvillei)
Sori pairs spaced along the frond
(Diplora sp.)
11a. Fronds coriaceous or subcoriaceous
12
Fronds herbaceous
17
12a. Fronds over 25 cm long
13
Fronds under 25 cm long
16
13a. Apex acute or acuminate
14
Apex cuspidate
A. werneri
14a. Surface squamiferous
A. vittaeforme
Surface naked
15
15a. Costa prominent
A. acrobryum
Costa depressed
A. comosum
16a. Rigidly coriaceous
A. durum
Subcoriaceous
A. morobense
17a. Rhizome slender, green
A. papuanum
Rhizome stouter, brown
A. amboinense
18a. Fronds +/- pinnatifid; sori in pairs facing each other
(Diplora translucens)
Fronds +/- pinnate or more compount; sori not paired and all facing +/- apically
19
19a. Fronds pinnate to bipinnatifid
20
Fronds bipinnate or more compound
41
20a. Small ferns with axes dark and polished
21
Mostly larger, axes not dark and polished
24
21a. Pinnae rounded at apex
22
Pinnae +/- acute
23
22a. Frond under 1 cm wide
A. trichomanes
Fronds broader
A. normale
23a. Apex of pinnae nearly equal-sided
A. unilaterale
Lower side of pinnae almost wanting
A. filipes
24a. Lamina herbaceous, green
25
Lamina more firm, usually brownish
32
25a. Pinnae large, not very oblique
26
Lower or all pinnae conspicuously oblique
27
26a. Rachis not winged
A. subemarginatum
Rachiswinged upward
A. regis
27a. Proliferous by slender rhizomes
A. bipinnatifidum
Not proliferous by rhizomes
28
28a. Pinnae under 5 cm long
29
Pinnae larger
31
29a. Lower pinnae distant, reduced
A. monotis
Frond truncate at base
30
30a. Pinnae merely toothed
A tenerum
Pinnae pinnatifid
A. decorum
31a. Surface naked
A. remotum
Surface squamulose
A. persicifolium
32a. Fronds up to c.5 cm wide
33
Well-developed fronds larger
36
33a. Larger pinnae incised
34
Pinnae not lobed
35
34a. Fronds up to 2.5 cm wide
A. gracile (A. clemensiae)
Fronds c.5 cm wide
A. planicaule
35a. Frond long and lax
A. pellucidum
Frond compact
A. Keysserianum
36a. Rachis proliferous
A. cromwellianum
Not proiferous
37
37a. Pinnae excised on lower side
38
Base incompletely excised
39
38a. Rachis +/- scaly
A. acutiusculum
Rachis glabrescent
A. falcatum
39a. Pinnae rather deeply lobed; sori at a very acute angle to costa
A. caudatum
Pinnae shallowly toothed or subentire; sori more divergent from coata
40
40a. Pinnae distinctly stalked
A. macrophyllum
Pinnae almost sessile
A. sanctichristophori
41a. Sori dorsal (not margial)
42
Sori elongate, straight, free side marginal
55
41c. Sori short, marginal to apical (Loxoscaphe)
56
42a. Segments linear-cueiform, 1 mm wide
A. insiticum
Segments distinctly wider
43
43a. Rachis bulbiferous
44
Not proliferous
45
44a. Pinnules toothed
A. hapalophyllum
Larger pinnules subpinnate
A. foersteri
44c. Tripinnate
A. praedigens
45a. Indusia with parallel sides
46
Indusia boat-shaped, ends acute
A. nutans
46a. Pinnules not deeply cut
47
Pinnules incised or pinnate
48
47a. Fronds clustered, thin
A. kelelense
Fronds remote, firm
A. setisectum
48a. Segmetns 2-3 mm wide
49
Segments broad
52
49a. Rhizome scandent
A. brassii
Rhizome short
50
50a. Rachis fibrillose
A. lanceolatum
Rachis glabrescent
51
51a. Frond lax
A. laxifolium
Frond compact
A. tafanum
52a. Fronds firm to coriaceous
53
Fronds thin
54
53a. Pinnules rounded
A. cuneatum
Pinnules subacute
A. affine
54a. Bright green
A. laserpitiifolium
Black green
A. shawii
55a. Frond pinnate or subbipinnate, small
A. bipinnatifidum
Frond decompound, large
A. scandens
56a. Fronds up to 10 cm long
L. bakeri
Fronds much larger
57
57a. Segments mostly entire
A. novoguineense
Fertile segments mostly furcate
A. schultzei

Diplora

Species in Papuasia

1a. Fronds deeply pinnatifid, subpinnate, lamina thin
D. translucens
Mature fronds simple (juvenile fronds may be finely divided), firm
2
2a. Fronds uniform, oblanceolate; rhizome compact
D. schizocarpa
Juvenile fronds compound, mature fronds linear-lanceolate; rhizome long-creeping
3
3a. Sori on most veins of fully fertile fronds
D. durvillei
Sori pairs spaced along the frond
Diplora sp.