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Asplenium
Key modified ex Copeland 1949, Philip, J. Sci. 78:207 (incl. Diplora) and Holttum 1974, Gard. Bull. Sing. 27:145 (sect. Thamnopteris).
Species in Papuasia
| 1a.
|
Fronds simple and entire |
2
|
| Fronds pinnatifid, pinnate or more compound |
18
|
|
| 2a. | Fronds with a continuous intramarginal vein, often nest-forming |
(sect. Thamopteris)
3 |
| Veins free to the margin, fronds not nest-forming |
8
|
|
| 3a. | Sori on almost all veins, including both branches of a vein which forks near costa; indusia less than 0.5 mm wide |
4
|
| Sori usually on one branch of each vein; indusia more than 0.5 mm wide |
7
|
|
| 4a. | Fronds nest-forming |
5
|
| Fronds not nest forming |
A. spirale
|
|
| 5a. | Fronds to almost 20 cm wide; apex gradually attenuate; costa prominent and squarish above, +/- flattened below |
A. nidus var. nidus
|
| Fronds 25-25 cm wide; apex broadly rounded; costa rounded above, rounded keeled below |
6
|
|
| 6a. | Fronds to c. 50 cm long, +/- ascending |
A. nidiforme
|
| Fronds 1m or more long +/- arching |
A. musifolium
|
|
| 7a. | Fronds to 20 cm or more wide with dilated bases; indusia more than 1 mm wide |
A. cymbifolium
|
| Fronds not over 12 cm wide, bases not dilated; indusia less than 1 mm wide |
A. phylitidis ssp. Malesicum
|
|
| 8a. | Sori paired, facing one another |
(Diplora)
9 |
| Sori not paired, all facing apically |
(Asplenium)
11 |
|
| 9a. | Fronds uniform, oblanceolate |
A. schizocarpa
|
| Juvenile fronds compound, mature fronds linear-lanceolate |
10
|
|
| 10a. | Sori on most veins of fully fertile fronds |
A. scolopendropsis (Diplora durvillei)
|
| Sori pairs spaced along the frond |
(Diplora sp.)
|
|
| 11a. | Fronds coriaceous or subcoriaceous |
12
|
| Fronds herbaceous |
17
|
|
| 12a. | Fronds over 25 cm long |
13
|
| Fronds under 25 cm long |
16
|
|
| 13a. | Apex acute or acuminate |
14
|
| Apex cuspidate |
A. werneri
|
|
| 14a. | Surface squamiferous |
A. vittaeforme
|
| Surface naked |
15
|
|
| 15a. | Costa prominent |
A. acrobryum
|
| Costa depressed |
A. comosum
|
|
| 16a. | Rigidly coriaceous |
A. durum
|
| Subcoriaceous |
A. morobense
|
|
| 17a. | Rhizome slender, green |
A. papuanum
|
| Rhizome stouter, brown |
A. amboinense
|
|
| 18a. | Fronds +/- pinnatifid; sori in pairs facing each other |
(Diplora translucens)
|
| Fronds +/- pinnate or more compount; sori not paired and all facing +/- apically |
19
|
|
| 19a. | Fronds pinnate to bipinnatifid |
20
|
| Fronds bipinnate or more compound |
41
|
|
| 20a. | Small ferns with axes dark and polished |
21
|
| Mostly larger, axes not dark and polished |
24
|
|
| 21a. | Pinnae rounded at apex |
22
|
| Pinnae +/- acute |
23
|
|
| 22a. | Frond under 1 cm wide |
A. trichomanes
|
| Fronds broader |
A. normale
|
|
| 23a. | Apex of pinnae nearly equal-sided |
A. unilaterale
|
| Lower side of pinnae almost wanting |
A. filipes
|
|
| 24a. | Lamina herbaceous, green |
25
|
| Lamina more firm, usually brownish |
32
|
|
| 25a. | Pinnae large, not very oblique |
26
|
| Lower or all pinnae conspicuously oblique |
27
|
|
| 26a. | Rachis not winged |
A. subemarginatum
|
| Rachiswinged upward |
A. regis
|
|
| 27a. | Proliferous by slender rhizomes |
A. bipinnatifidum
|
| Not proliferous by rhizomes |
28
|
|
| 28a. | Pinnae under 5 cm long |
29
|
| Pinnae larger |
31
|
|
| 29a. | Lower pinnae distant, reduced |
A. monotis
|
| Frond truncate at base |
30
|
|
| 30a. | Pinnae merely toothed |
A tenerum
|
| Pinnae pinnatifid |
A. decorum
|
|
| 31a. | Surface naked |
A. remotum
|
| Surface squamulose |
A. persicifolium
|
|
| 32a. | Fronds up to c.5 cm wide |
33
|
| Well-developed fronds larger |
36
|
|
| 33a. | Larger pinnae incised |
34
|
| Pinnae not lobed |
35
|
|
| 34a. | Fronds up to 2.5 cm wide |
A. gracile (A. clemensiae)
|
| Fronds c.5 cm wide |
A. planicaule
|
|
| 35a. | Frond long and lax |
A. pellucidum
|
| Frond compact |
A. Keysserianum
|
|
| 36a. | Rachis proliferous |
A. cromwellianum
|
| Not proiferous |
37
|
|
| 37a. | Pinnae excised on lower side |
38
|
| Base incompletely excised |
39
|
|
| 38a. | Rachis +/- scaly |
A. acutiusculum
|
| Rachis glabrescent |
A. falcatum
|
|
| 39a. | Pinnae rather deeply lobed; sori at a very acute angle to costa |
A. caudatum
|
| Pinnae shallowly toothed or subentire; sori more divergent from coata |
40
|
|
| 40a. | Pinnae distinctly stalked |
A. macrophyllum
|
| Pinnae almost sessile |
A. sanctichristophori
|
|
| 41a. | Sori dorsal (not margial) |
42
|
| Sori elongate, straight, free side marginal |
55
|
|
| 41c. | Sori short, marginal to apical (Loxoscaphe) |
56
|
| 42a. | Segments linear-cueiform, 1 mm wide |
A. insiticum
|
| Segments distinctly wider |
43
|
|
| 43a. | Rachis bulbiferous |
44
|
| Not proliferous |
45
|
|
| 44a. | Pinnules toothed |
A. hapalophyllum
|
| Larger pinnules subpinnate |
A. foersteri
|
|
| 44c. | Tripinnate |
A. praedigens
|
| 45a. | Indusia with parallel sides |
46
|
| Indusia boat-shaped, ends acute |
A. nutans
|
|
| 46a. | Pinnules not deeply cut |
47
|
| Pinnules incised or pinnate |
48
|
|
| 47a. | Fronds clustered, thin |
A. kelelense
|
| Fronds remote, firm |
A. setisectum
|
|
| 48a. | Segmetns 2-3 mm wide |
49
|
| Segments broad |
52
|
|
| 49a. | Rhizome scandent |
A. brassii
|
| Rhizome short |
50
|
|
| 50a. | Rachis fibrillose |
A. lanceolatum
|
| Rachis glabrescent |
51
|
|
| 51a. | Frond lax |
A. laxifolium
|
| Frond compact |
A. tafanum
|
|
| 52a. | Fronds firm to coriaceous |
53
|
| Fronds thin |
54
|
|
| 53a. | Pinnules rounded |
A. cuneatum
|
| Pinnules subacute |
A. affine
|
|
| 54a. | Bright green |
A. laserpitiifolium
|
| Black green |
A. shawii
|
|
| 55a. | Frond pinnate or subbipinnate, small |
A. bipinnatifidum
|
| Frond decompound, large |
A. scandens
|
|
| 56a. | Fronds up to 10 cm long |
L. bakeri
|
| Fronds much larger |
57
|
|
| 57a. | Segments mostly entire |
A. novoguineense
|
| Fertile segments mostly furcate |
A. schultzei
|
Diplora
Species in Papuasia
| 1a. | Fronds deeply pinnatifid, subpinnate, lamina thin |
D. translucens
|
| Mature fronds simple (juvenile fronds may be finely divided), firm |
2
|
|
| 2a. | Fronds uniform, oblanceolate; rhizome compact |
D. schizocarpa
|
| Juvenile fronds compound, mature fronds linear-lanceolate; rhizome long-creeping |
3
|
|
| 3a. | Sori on most veins of fully fertile fronds |
D. durvillei
|
| Sori pairs spaced along the frond |
Diplora sp.
|