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Sphaerostephanos J.Sm.

Terrestrial ferns of small to large size. Rhizome a short-creeping or compact erect caudex, sometimes arborescent, rarely long-creeping, radially symmetric, dictyostelic, bearing narrow, thin scales with marginal and superficial unicellular, acicular hairs. Fronds long-stipitate, the stipe with 2 vascular bundles at the base uniting upward into a single U-shaped vascular strand, lamina pinnate with +/- entire, crenate or pinnately deeply lobed pinnae, margins of lobes entire, basal pinnae much reduced, usually abruptly but sometimes gradually reduced, rarely not reduced, the apex of frond rarely pinna-like, swollen aerophores often present at bases of pinnae, midribs of pinnae grooved above, but the grooves not confluent with rachis groove, costae and costules of both surfaces with acicular hairs, often also between the veins, yellow, sessile, sperical, glands, often present on one or both surfaces, lower surface rarely pustular when dry, the veins in a pinnate group in each pinna-lobe or crenation (if present), the basal pairs in deeply lobed species, or all veins in crenate or entire species, anastomosing to form an excurrent vein terminating in the sinus membrane, the remainder free, simple, terminating in the margin, or all veins free. Sori indusiate, less often exindusiate, round or +/- elongate along the veins, +/- medial on the veins (not excurrent veins), indusium rather thin, often hairy or glandular or both; sporangia on a 2-seriate stalk bearing a stalked, glandular hair, body of sporangia often with short acicular hairs or yellow, sessile, spherical glands borne either side of the annulus, the annulus longitudinal, interrupted. Spores monolete, mostly pale brown, spinulose, or with many small wings, or with a continuous wing and many cross-wings.

Key modified ex Holttum 1981, F1. Males. ser. 2, 1:439 and 1977, Allertonia 1:202.