D. Christine Cargill

Curator of cryptogam collections

Ph.D, Southern Illinois University, Illinois, USA
B.Sc. (Hons.) Monash University, Melbourne, Australia

Photo of Christine Cargill

Phone (+61) 02 6250 9464


Centre for Australian National Biodiversity Research
CSIRO Plant Industry
GPO Box 1700
Canberra ACT 2601


Systematics, phylogeny and biogeography of the Australasian liverworts and hornworts, in particular the families Aytoniaceae, Fossombroniaceae, Ricciaceae and Riellaceae.


Dawes, T.N, Villarreal, J.C., Szövéyi, P., Bisang, I., Li, F-W., Hauser, D.A., Quandt, D., Cargill, D.C. & Forrest, L.L. (2020). Extremely low genetic diversity in the European clade of the model bryophyte Anthoceros agrestis. Plant Systematics and Evolution. 306(2): 1–10.

Cargill, D.C., Manju C. N., Deepa K. M., Chandini V. K. & Rajesh K. P. (2019). A new Indian species of Riccia L. with connections to northern Australia, Journal of Bryology. 41(3): 236–242.

Lepschi, B.J., Cargill, D.C., Albrecht, D.E. & Monro, A.M. (eds) (30 August 2019). Census of the Flora of the Australian Capital Territory. Version 4.1.

Söderström, L., A.Hagborg, M. von Konrat, S.Bartholomew-Began, D.Bell, L.Briscoe, E.Brown, D.C.Cargill, D.P.Costa, B.J.Crandall-Stotler, E.D.Cooper, G.Dauphin, J.J.Engel, K.Feldberg, D.Glenny, S.R.Gradstein, X.He, A.L.Ilkiu-Borges, J.Heinrichs, J.Hentschel, T.Katagiri, N.A.Konstantinova, J.Larraín, D.G.Long, M.Nebel, T.Pócs, F.Puche, M.E.Reiner-Drehwald, M.A.M.Renner, A.Sass-Gyarmati, A.Schäfer-Verwimp, J.G.Segarra Moragues, R.E.Stotler, P.Sukkharak, B.M.Thiers, J.Uribe, J.Váňa, J.C.Villarreal, M.Wigginton, L.Zhang, & R.-L.Zhu (2016). World checklist of hornworts and liverworts. PhytoKeys 59: 1–828.

Nagalingum, N.S., N.Knerr, B.D.Mishler & D.C.Cargill (2015). Overlapping fern and bryophyte hotspots: Assessing ferns as a predictor of bryophyte diversity. Telopea 17: 383–392.

Villarreal, J.C., D.C.Cargill, L.Söderström, A.Hagborg, & M. von Konrat (2015). Notes on Early Land Plants Today. 70. Nomenclatural notes in hornworts (Anthocerotophyta). Phytotaxa 208: 92–96.

Gonzalez-Orozco, C.E., M.C.Ebach, S.Laffan, A.H.Thornhill, N.J.Knerr, A.N.Schmidt-Lebuhn, D.C.Cargill, M.Clements, N.S.Nagalingum, B.D.Mishler & J.T.Miller (March 2014). Quantifying phytogeographical regions of Australia using geospatial turnover in species composition. PLOS ONE. 9(3): e92558.

Cargill, D.C., N.G.F.Vella, I.Sharma & J.T.Miller (2013). Cryptic speciation and species diversity among Australian and New Zealand hornwort taxa of Megaceros Campb. (Dendrocerotaceae). Australian Systematic Botany. 26(5): 356–377.

Cargill, D.C. & J.Milne (2013). A new terrestrial genus and species within the aquatic liverwort family Riellaceae (Sphaerocarpales) from Australia. Polish Botanical Journal 58(1): 71–80.

Cargill, D.C., L.Söderström, A. Hagborg & M.von Konrat (2013). Notes on Early Land Plants Today. 23. A new synonym in Anthoceros (Anthocerotaceae, Anthocerotophyta). Phytotaxa 76(3): 48–49.

Cargill, D.C., L.Söderström, A.Hagborg & M.von Konrat (2012). Notes on Early Land Plants Today. 3. Fossombronia ruminata Cargill, a new name for Fossombronia maritima G.A.M. Scott et D.C. Pike ex Cargill, nom. illeg. Phytotaxa 65: 45.

Stevenson, L.A., C.E.Gonzalez-Orozco, N.Knerr, D.C.Cargill & J.T.Miller (2012). Species richness and endemism of Australian bryophytes. Journal of Bryology 34(2): 101–107.

Milner,M., I.Sharma, V.Freire & D.C.Cargill (2011). Spore ornamentation patterns and species delimitations within Australian Fossombronia Raddi (Fossombroniaceae) populations. The Bryologist 114: 114: 128–141.

Villarreal, J.C., W.Frey & D.C.Cargill (eds) (2010). Bryophyte Biology, Phylogeography, Systematics and Evolution in the Southern Hemisphere: A volume in honour of Dr. Gabriela Hässel de Menéndez†. Nova Hedwigia 91.

Villarreal, J.C., D.C.Cargill & B.Goffinet (2010). Phaeomegaceros squamuligerus subspecies hasselii (Dendrocerotaceae), a new taxon from the Southern Hemisphere. Nova Hedwigia 91: 349–360.

Villarreal, J. C., D. C. Cargill, A. Hagborg, L. Söderström & R. S. Renzaglia (2010). A synthesis of hornwort diversity: Patterns, causes and future work. Phytotaxa 9: 150–166.

Villarreal, J.C., B.Goffinet, R.J.Duff & D.C.Cargill (2010). Phylogenetic delineation of Nothoceros and Megaceros (Dendrocerotaceae). The Bryologist 113(1): 106–113.

Cargill, D. C. & B. A. Fuhrer (2008). Taxonomic studies of the Australian Anthocerotophyta II: The genus Phaeoceros. Fieldiana, Botany, n.s. 47: 239–253.


  • Taxonomy, systematics and phylogeny of Australian hornworts.
  • Taxonomy and phylogeny of Australian members of the Ricciaceae
  • Riccias of the Northern Territory
  • Revision of Australian members of the Riellaceae

  • Updated 18 June, 2020 , webmaster, CANBR (