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Australian Biological Resources Study

Checklist of the Lichens of Australia and its Island Territories
Introduction | A–D | E–O | P–R | S–Z | Oceanic Islands | References
Ocellularia papillata (Leight.) Zahlbr.

Cat. Lich. Univ. 2: 597 (1923)

Thelotrema papillatum Leight., Trans. Linn. Soc. London 27: 169 (1869).

T: Central Province, Ceylon [Sri Lanka], G.H.K.Thwaites (CL 129); lecto: BM, fide M.E.Hale, Smithsonian Contr. Bot. 16: 24 (1974); isolecto: H-NYL 3854, S, UPS.

Thelotrema collativum Kremp., Nuovo Giorn. Bot. Ital. 7: 20 (1875); Ocellularia collativa (Kremp.) Zahlbr., Cat. Lich. Univ. 2: 587 (1923). T: Sarawak, [Malaysia], O.Beccari 39b; holo: M.

Ocellularia viridipallens Müll.Arg., Flora 70: 397 (1887). T: Trinity Bay, [Cairns], Qld, W.A.Sayer s.n.; holo: G.

Thelotrema zamboangense Vain., Ann. Acad. Sci. Fenn., ser. A, 15(6): 192 (1921); Ocellularia zamboangensis (Vain.) Zahlbr., Cat. Lich. Univ. 2: 604 (1923). T: Zamboanga, Mindanao, Philippines, E.D.Merrill 8342; iso: US.

Ocellularia ascidioidea Hale, Bull. Brit. Mus. (Nat. Hist.), Bot. 8: 300 (1981). T: Southern Province, Sri Lanka, M.E.Hale 47035; holo: US.


  Thallus endophloeodal to epiphloeodal, to c. 200 µm thick, pale greenish grey to olive or brownish, dull to glossy, smooth, continuous to verrucose or verruculose, rimose or not. True cortex ±continuous, to c. 50 µm thick, formed by irregular hyphae. Algal layer well developed, continuous; calcium oxalate crystals small to moderately large, scattered or clustered. Ascomata ±conspicuous, apothecioid at maturity, to c. 0.8 mm diam., ±rounded to irregular, solitary to fused, immersed to moderately emergent, often verrucose, hemispherical to urceolate. Disc with the columella visible in mature ascomata, ±distinctly pruinose, off-white to brownish or dark grey, often fused with the thalline rim, entire. Pores formed by the thalline rim margin, c. 0.4 mm diam., ±rounded to irregular, thin to thick, occasionally depressed, entire to split and/or evanescent, brownish to dark grey due to the protuberant proper exciple, otherwise concolorous with the thallus; thalline rim incurved. Proper exciple fused, thin, yellowish brown to brownish internally, dark brown or carbonised marginally and distally, often amyloid at the base. Hymenium to c. 150 µm thick, not inspersed, distinctly conglutinated; paraphyses straight to bent, slightly interwoven, unbranched, with slightly thickened tips; columellar structures well developed in older ascomata, to 300 µm wide, entire, brownish to carbonised, with or without a ±thick covering layer of greyish granules. Epihymenium hyaline, sometimes with greyish or brownish granules. Asci 8-spored; tholus initially thick, thin when mature. Ascospores transversely septate, oblong to fusiform or ellipsoidal, with ±rounded to narrowly rounded or rarely subacute ends, hyaline, distinctly amyloid, 10–35 (–45) × 6–10 µm, with 4–10 locules; locules initially ±rounded to slightly angular, then ±lentiform to oblong or rectangular, with hemispherical to conical end cells; septa moderately thick, regular; ascospore wall and endospore moderately thick; wall non-halonate. Pycnidia not seen.
CHEMISTRY: Thallus K–, C–, P–; no compounds detectable by TLC.
  Corticolous in forest in eastern Qld and N.S.W.; mainly pantropical.  
  Mangold et al. (2009)  

Checklist Index
Introduction | A–D | E–O | P–R | S–Z | Oceanic Islands | References

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